I re-read what I wrote, and I said intake charge...sorry thats not what I meant. When you're dealing with resonant tuning, you're dealing with sonic speeds. The actual intake charge, no. But the wave produced by the valve slamming shut, it at sonic speeds. That wave is what is used to tune the runner lengths to provide passive supercharging. The intake charge comes in, the valve slams shut. There is a sonic wave produced, and at the same time, the intake charge compresses behind the intake valve. The sonic wave reflects back toward the plenum. After the air charge compresses, it reflects back toward the plenum, and in a sense, it stretches and then bounces off the pressure of the plenum...kind of like a pillow. Think of the plenum volume as being a pillow of sorts. Too much volume will cause the returning wave to be dampened...just like it would be with a big pillow. Too little volume, the reflective power will be high, but plenum volume will be too small to provide the balanced air supply the runners need. There are 2 waves moving at different speeds. When you start calculating runner lengths, the formulas are based on timing resonating sonic waves which help to push the resonating intake charge back into the intake valve the next time it opens, The air speeds are more or less a function of a pressure differential, (air density plays in here) the runner cross sectional area, piston acceleration, cam timing...a whole bunch of things. The old school of thought with forced induction and runner lengths, was that tuned runner lengths were no big deal when under boost. Turns out to not be the case at all. Resonant tuning will provide the same benefits in boosted applications. The runner lengths on these cars are tuned for very high rpm. The torque curve would be much stronger with longer runners, (about 12" for a peak hp at 6000rpm) but they still do fine. These motors have to rely on sheer airflow and doesnt really use resonant tuning to boost the torque curve.
Ive done a huge amount of work with this stuff in the past, and the difference runner lengths make is astounding. I used my old car as a testbed, and tried about 20 different variations of runner lengths, plenum volumes, etc. Based on the TR runner length, if it were ever built in N/A form, the TR runner length would be terrible for providing a useful torque curve. On my N/A 99 cobra, between a 12" runner and a 7 inch runner, there was about 100 lb ft of torque lost at about 3000rpm. Peak torque was down about 50lbs. Peak hp increased about 8 hp, but the engine didnt even reach the stock hp curve until about 6000rpm, and then peaked at 7000, vs. 6000 before. The car was a DOG. This is just one out of about 20 different combos.